Layered entrenchment maintains essentiality in the evolution of Form I Rubisco complexes
2024

How Rubisco Became Dependent on a Small Subunit

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Author Information

Author(s): Luca Schulz, Jan Zarzycki, Wieland Steinchen, Georg K A Hochberg, Tobias J Erb

Primary Institution: Max Planck Institute for Terrestrial Microbiology

Hypothesis

What factors cause essentiality to persist over evolutionary timescales in protein complexes?

Conclusion

Rubisco's dependence on its small subunit has evolved through multiple, genetically distinct mechanisms that ensure its essentiality over billions of years.

Supporting Evidence

  • Rubisco's essentiality can arise through the accumulation of changes that are tolerated in the complex state but would be harmful for standalone components.
  • Layering multiple mechanisms of essentiality can lead to its persistence, even if any given mechanism reverts.
  • New interaction partners can drastically reshape which substitutions are tolerated in the proteins they are recruited into.

Takeaway

Rubisco, a key enzyme in photosynthesis, needs a small partner to work properly, and this need developed in different ways over a very long time.

Methodology

The study used ancestral sequence reconstruction and various biochemical assays to investigate the mechanisms behind Rubisco's dependence on its small subunit.

Digital Object Identifier (DOI)

10.1038/s44318-024-00311-1

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